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Faculty & Staff Profile
Marine Science Center 205
207-602-2410 (w)
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Stephan I. Zeeman, PhD
Professor, Department of Marine Sciences


Oceanography, phytoplankton, primary production, remote sensing, geographic information systems, Bering Sea ecosystems.


A.A., Biology, Pasadena City College

B.A., Environmental Biology, University of California, Santa Barbara

M.S., Botany, University of Wisconsin, Milwaukee,

Ph.D., Marine Science, University of South Carolina, Columbia

Post-Grad Training

Post-Doctoral Training
Phytoplankton Physiology, Harbor Branch Oceanographic Institute, Fort Pierce, Florida

Research and Scholarship

Research Interests

Phytoplankton, primary production, remote sensing, GIS, satellite tracking, food web dynamics, habitat selection by marine animals (whales, basking sharks), K-12 science education

Current Research or Scholarship

Land-Use changes and their effects on pathogens and nutrient transport through rivers to coastal waters.  Migration and habitat selection by basking sharks. Photosynthesis as measured by Pulse Amplitude Modulated Fluorometry. Physical-biological coupling of primary production in the Bering Sea.

Selected Publications

Skomal,G.B., S.I. Zeeman, J.H. Chisholm, E.L. Summers, H.J. Walsh, K.W. McMahon, and S.R. Thorrold. 2009. Transequatorial Migrations by Basking Sharks in the Western Atlantic Ocean, Current Biology, doi:10.1016/j.cub.2009.04.019

Raymond N. Sambrotto, Calvin Mordy, Stephan I. Zeeman, Phyllis J. Stabeno and S. Allen Macklin. 2008. Physical forcing and nutrient conditions associated with patterns of Chl a and phytoplankton productivity in the southeastern Bering Sea during summer.Deep Sea Research Part II: Topical Studies in Oceanography. 55:1745-1760.

Summers, Erin L., James A. Estrada and Stephan I. Zeeman. 2006 A note on geographic and seasonal fluctuations in the isotopic composition of baleen in four North Atlantic right whales (Eubalaena glacialis). J. Cetacean Res. Manage. 8(3):241-245;  

Jahncke, J., K. O. Coyle, S. I. Zeeman, N. B. Kachel, G. L. Hunt Jr. 2005. Distribution of foraging shearwaters relative to inner front of SE Bering Sea. Mar. Ecol. Prog. Ser. 305: 219-233;  

Mordy,Calvin W., Phyllis J. Stabeno, Carol Ladd, Stephan Zeeman, David P. Wisegarver, Sigrid A. Salo and George L. Hunt, Jr. 2005. Nutrients and primary production along the eastern Aleutian Island Archipelago. Fish. Oceanogr. 14 (Suppl. 1), 55-76;

A. Merico, T. Tyrrell, E.J. Lessard, T. Oguz, P.J. Stabeno, S.I. Zeeman, and T.E. Whitledge. 2004. Modelling the Bering Sea shelf ecosystem: role of climate influences and trophic interactions in generating Emiliania huxleyi blooms 1997-2000. Deep Sea Res. I 51: 1803-1826.  

Recently Funded Grants

2009, The Interactions of Biology, Chemistry and Physics at the Land-Ocean Interface: A Systemic PARTnership Aimed at Connecting University and School (SPARTACUS).NSF DGE-0841361, $3.78M, 5 yrs.

Joint NASA Center for Remote Sensing at the University of New England (2003) NASA NAG13-03027, $581,000, 1 year. 

Collaborative Proposal: Cross-Shelf Transport and Post-Bloom New Production Near the Pribilof Islands (2003). NSF OPP-0326415, $128,479, 2 years. 

Students and Scientists: Together Advancing Science Knowledge (2002) NSF ESI-0222740, $350,000, 3 1/2 years. Saco Bay Scallop Stock Enhancement Project, Northeast Consortium (2002), $25,000, 1 year.

Student Opportunities: Yes