Stephan I. Zeeman



University of South Carolina, Columbia



University of Wisconsin, Milwaukee,



University of California, Santa Barbara



Pasadena City College


Post-Doctoral Training

Phytoplankton Physiology

Harbor Branch Oceanographic Institute

Fort Pierce





primary production

remote sensing

geographic information systems

Bering Sea ecosystems.


Current Research

Land-Use changes and their effects on pathogens and nutrient transport through rivers to coastal waters. Migration and habitat selection by basking sharks. Photosynthesis as measured by Pulse Amplitude Modulated Fluorometry. Physical-biological coupling of primary production in the Bering Sea.

Research Interests

Phytoplankton ecology, GIS, Remote Sensing

Selected Publications


Goodchild, C.G., Frederich, M., Zeeman, S.I.  2015. Is altered behavior linked to cellular energy regulation in a freshwater mussel (Elliptio complanata) exposed to triclosan? Comparative Biochemistry and Physiology- Part C: (179:150-157).


Zeeman, S.I. and T. Spillane. 2015. Water Quality in the Saco River. Chapter 10 pp. 113-128 In: Sustaining the Saco Estuary, Final Report,  part of Maine's Sustainability Solutions Initiative . 128 pages.


Goodchild,C.G., M. Frederich, S.I. Zeeman. 2015. AMP-activated protein kinase is a biomarker of energetic status in freshwater mussels exposed to municipal effluents. Science of the Total Environment. 512-513, pp. 201-209


Curtis , Tobey H., Stephan I Zeeman , Erin L Summers , Steven X Cadrin and Gregory B Skomal. 2014. Eyes in the sky: linking satellite oceanography and biotelemetry to explore habitat selection by
basking sharks.   Animal Biotelemetry 2014, 2:12 DOI: 10.1186/2050-3385-2-12


Susan J. Hillman , Kylie H. Bloodsworth , Charles E. Tilburg , Stephan I. Zeeman  & Henrietta E. List. 2013.  K-12 Students' Perceptions of Scientists: Finding a valid measurement and exploring whether exposure to scientists makes an impact.   International Journal of Science Education, 36:15, 2580-2595, DOI: 10.1080/09500693.2014.908264


Zeeman, S.I. & P.Weinstein (Author/editors). 2012 Chapter 3, Water, water quality and health. In: Environmental Tracking for Public Health Surveillance, S.A. Morain and A. Budge (Eds.),ISPRS Book Series Vol. 11. CRC Press, 480 Pages


Tilburg, Charles E.,  Shaun M. Gill, Stephan I. Zeeman, Amy E. Carlson, Timothy W. Arienti, Jessica A. Eickhorst, and Philip O. Yund. 2011. Characteristics of a Shallow River Plume: Observations
from the Saco River Coastal Observing System.  Estuaries and Coasts 34:785–799
DOI 10.1007/s12237-011-9401-y


Skomal,G.B., S.I. Zeeman, J.H. Chisholm, E.L. Summers, H.J. Walsh, K.W. McMahon, and S.R. Thorrold. 2009. Transequatorial Migrations by Basking Sharks in the Western Atlantic Ocean, Current Biology, doi:10.1016/j.cub.2009.04.019


Sambrotto, Raymond N., Calvin Mordy, Stephan I. Zeeman, Phyllis J. Stabeno and S. Allen Macklin. 2008. Physical forcing and nutrient conditions associated with patterns of Chl a and phytoplankton productivity in the southeastern Bering Sea during summer.Deep Sea Research Part II: Topical Studies in Oceanography. 55:1745-1760.


Summers, Erin L., James A. Estrada and Stephan I. Zeeman. 2006 A note on geographic and seasonal fluctuations in the isotopic composition of baleen in four North Atlantic right whales (Eubalaena glacialis). J. Cetacean Res. Manage. 8(3):241-245;  


Jahncke, J., K. O. Coyle, S. I. Zeeman, N. B. Kachel, G. L. Hunt Jr. 2005. Distribution of foraging shearwaters relative to inner front of SE Bering Sea. Mar. Ecol. Prog. Ser. 305: 219-233;  


Mordy,Calvin W., Phyllis J. Stabeno, Carol Ladd, Stephan Zeeman, David P. Wisegarver, Sigrid A. Salo and George L. Hunt, Jr. 2005. Nutrients and primary production along the eastern Aleutian Island Archipelago. Fish. Oceanogr. 14 (Suppl. 1), 55-76;


Merico, A., T. Tyrrell, E.J. Lessard, T. Oguz, P.J. Stabeno, S.I. Zeeman, and T.E. Whitledge. 2004. Modelling the Bering Sea shelf ecosystem: role of climate influences and trophic interactions in generating Emiliania huxleyi blooms 1997-2000. Deep Sea Res. I 51: 1803-1826.  

Other Scholarly Activity

Fulbright Fellowship to travel to Indonesia 2014-2015.

Funded Grants

2014.  TURBO: The Undergraduate Saco River Biodiversity Observatory -- A Long-Term Ecological Research-Style Research Experience to Enhance STEM Education, Co-PI,  NSF DUE-1431955, $639,246, 5 yrs.

2013. MRI: Acquisition of a Gas CHromatograhy-Mass SPectrometry Instrument for Research and Teaching.  NSF MRI-1229519, $137,527, 2 yrs.

2009, The Interactions of Biology, Chemistry and Physics at the Land-Ocean Interface: A Systemic PARTnership Aimed at Connecting University and School (SPARTACUS).NSF DGE-0841361, $3.78M, 5 yrs.

Joint NASA Center for Remote Sensing at the University of New England (2003) NASA NAG13-03027, $581,000, 1 year.

Collaborative Proposal: Cross-Shelf Transport and Post-Bloom New Production Near the Pribilof Islands (2003). NSF OPP-0326415, $128,479, 2 years.

Students and Scientists: Together Advancing Science Knowledge (2002) NSF ESI-0222740, $350,000, 3 1/2 years. Saco Bay Scallop Stock Enhancement Project, Northeast Consortium (2002), $25,000, 1 year.

Research Topics

Professor Stephan I. Zeeman

Stephan I. Zeeman



Professor, Department of Marine Sciences

Marine Science Center

(207) 602-2410