Professor Stephan I. Zeeman

Stephan I. Zeeman, Ph.D.

Professor Emeritus

Professor Zeeman was born in Indonesia. During his youth his family moved to the Netherlands before coming to the United States. He lived and went to school in southern California. In college, he wanted to go into forestry but became fascinated with aquatic systems. As an undergraduate at UC Santa Barbara he conducted research on phytoplankton in sub-alpine lakes in the Sierra Nevada Mountains. He continued by studying phytoplankton in Lake Michigan for his M.S. at UW-Milwaukee, before turning to marine phytoplankton for his Ph.D. at the University of South Carolina. He conducted postdoctoral studies at Harbor Branch Oceanographic Institution in Florida. Here he became engaged in studies of the Bering Sea and Arctic Ocean in collaborative work with scientists from across the US and the Soviet Union. This was followed by a stint as Director of an environmental education center in South Florida. Dr. Zeeman joined the faculty at UNE in 1987 and continued phytoplankton research in the Bering Sea related to food webs.

Fulbright Fellow 2014-2015 in Indonesia

Hobbies: photography, hiking and camping.



University of South Carolina, Columbia
University of Wisconsin, Milwaukee,
University of California, Santa Barbara
Pasadena City College


Aquatic ecology
Geographic information systems
Marine ecology
Remote sensing

Post-Doctoral Training

Post-Doctoral Training, Phytoplankton Physiology
Harbor Branch Oceanographic Institute (Fort Pierce, Florida)


Current research

Land-Use changes and their effects on pathogens and nutrient transport through rivers to coastal waters. Migration and habitat selection by basking sharks. Photosynthesis as measured by Pulse Amplitude Modulated Fluorometry. Physical-biological coupling of primary production in the Bering Sea.

Selected publications

Liu, C.L., Zhai, L., Zeeman, S.I., (...), Moran, S.B., Lomas, M.W. 2016. Seasonal and geographic variations in modeled primary production and phytoplankton losses from the mixed layer between warm and cold years on the eastern Bering Sea shelf. Deep-Sea Research Part II: Topical Studies in Oceanography 134, pp. 141-156


Hillman, S.J., Zeeman, S.I., Tilburg, C.E., List, H.E. 2016. My Attitudes Toward Science (MATS): the development of a multidimensional instrument measuring students’ science attitudes. Learning Environments Research 19(2), pp. 203-219


Goodchild, C.G., Frederich, M., Zeeman, S.I.  2015. Is altered behavior linked to cellular energy regulation in a freshwater mussel (Elliptio complanata) exposed to triclosan? Comparative Biochemistry and Physiology- Part C: (179:150-157).

Zeeman, S.I. and T. Spillane. 2015. Water Quality in the Saco River. Chapter 10 pp. 113-128 In: Sustaining the Saco Estuary, Final Report,  part of Maine's Sustainability Solutions Initiative . 128 pages.

Goodchild,C.G., M. Frederich, S.I. Zeeman. 2015. AMP-activated protein kinase is a biomarker of energetic status in freshwater mussels exposed to municipal effluents. Science of the Total Environment. 512-513, pp. 201-209

Curtis , Tobey H., Stephan I Zeeman , Erin L Summers , Steven X Cadrin and Gregory B Skomal. 2014. Eyes in the sky: linking satellite oceanography and biotelemetry to explore habitat selection by
basking sharks.   Animal Biotelemetry 2014, 2:12 DOI: 10.1186/2050-3385-2-12

Susan J. Hillman , Kylie H. Bloodsworth , Charles E. Tilburg , Stephan I. Zeeman  & Henrietta E. List. 2013.  K-12 Students' Perceptions of Scientists: Finding a valid measurement and exploring whether exposure to scientists makes an impact.   International Journal of Science Education, 36:15, 2580-2595, DOI: 10.1080/09500693.2014.908264

Zeeman, S.I. & P.Weinstein (Author/editors). 2012 Chapter 3, Water, water quality and health. In: Environmental Tracking for Public Health Surveillance, S.A. Morain and A. Budge (Eds.),ISPRS Book Series Vol. 11. CRC Press, 480 Pages

Tilburg, Charles E.,  Shaun M. Gill, Stephan I. Zeeman, Amy E. Carlson, Timothy W. Arienti, Jessica A. Eickhorst, and Philip O. Yund. 2011. Characteristics of a Shallow River Plume: Observations
from the Saco River Coastal Observing System.  Estuaries and Coasts 34:785–799
DOI 10.1007/s12237-011-9401-y

Skomal,G.B., S.I. Zeeman, J.H. Chisholm, E.L. Summers, H.J. Walsh, K.W. McMahon, and S.R. Thorrold. 2009. Transequatorial Migrations by Basking Sharks in the Western Atlantic Ocean, Current Biology, doi:10.1016/j.cub.2009.04.019

Sambrotto, Raymond N., Calvin Mordy, Stephan I. Zeeman, Phyllis J. Stabeno and S. Allen Macklin. 2008. Physical forcing and nutrient conditions associated with patterns of Chl a and phytoplankton productivity in the southeastern Bering Sea during summer.Deep Sea Research Part II: Topical Studies in Oceanography. 55:1745-1760.

Summers, Erin L., James A. Estrada and Stephan I. Zeeman. 2006 A note on geographic and seasonal fluctuations in the isotopic composition of baleen in four North Atlantic right whales (Eubalaena glacialis). J. Cetacean Res. Manage. 8(3):241-245;  

Jahncke, J., K. O. Coyle, S. I. Zeeman, N. B. Kachel, G. L. Hunt Jr. 2005. Distribution of foraging shearwaters relative to inner front of SE Bering Sea. Mar. Ecol. Prog. Ser. 305: 219-233;  

Mordy,Calvin W., Phyllis J. Stabeno, Carol Ladd, Stephan Zeeman, David P. Wisegarver, Sigrid A. Salo and George L. Hunt, Jr. 2005. Nutrients and primary production along the eastern Aleutian Island Archipelago. Fish. Oceanogr. 14 (Suppl. 1), 55-76;

Merico, A., T. Tyrrell, E.J. Lessard, T. Oguz, P.J. Stabeno, S.I. Zeeman, and T.E. Whitledge. 2004. Modelling the Bering Sea shelf ecosystem: role of climate influences and trophic interactions in generating Emiliania huxleyi blooms 1997-2000. Deep Sea Res. I 51: 1803-1826.  

Other scholarly activity

Fulbright Fellowship to travel to Indonesia 2014-2015.

Research interests

Phytoplankton ecology, GIS, Remote Sensing, Robotics

Research topics

Biological Oceanography
Climate Change
Fisheries Ecosystems
K-12 Education
Marine Conservation Ecology
Marine Physiological Ecology
Salt Marsh Ecology